Wenzel , M A , Webster , L M I , Paterson , S , Mougeot , F , Martinez-Padilla , J & Piertney , S B 2013 , ' A transcriptomic investigation of handicap models in sexual selection ' Behavioral Ecology and Sociobiology , vol. 67 , no. 2 , pp. 221-234 . https://doi.org/10.1007/s00265-012-1442-0