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Acknowledgements A full list of acknowledgments appears in the Supplementary Note. Part of this work was conducted using the UK Biobank Resource under application number 9161.131166935engSDG 3 - Good Health and Well-beingQH426 GeneticsSupplementary DataQH426Journal article10.1038/s41588-018-0084-1https://www.nature.com/authors/policies/license.html504